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This chapter deals with the biology of squashes, pumkins, zucchinis and gourds (Cucurbita species). It contains information for use during the risk/safety regulatory assessment of genetically engineered varieties intended to be grown in the environment (biosafety). It includes elements of taxonomy, centres of origin and distribution, morphological characters, reproductive biology, genetics, hybridisation and introgression, crop production and cultivation practices, interactions with other organisms, pests and pathogens, and biotechnological developments.